Área de Brodmann 41

Cerebro: Área de Brodmann 41 áreas de Brodmann 41 & 42 of the human brain. The Primary Auditory Cortex is highlighted in magenta, and has been known to interact with all areas highlighted on this neural map. Sujeto latino de Gray # Part of Components Artery Vein BrainInfo/UW ancil-428 MeSH [1] The primary auditory cortex is the region of the brain that is responsible for processing of auditory (sonido) information. Contents 1 Function of primary auditory cortex 2 Área de Brodmann 41 3 Área de Brodmann 42 4 Relationship to auditory system 5 Referencias 6 Enlaces externos 7 See also Function of primary auditory cortex As with other primary sensory cortical areas, auditory sensations reach perception only if received and processed by a cortical area. Neurons in the auditory cortex are organised according to the frequency of sound to which they respond best. Neurons at one end of the auditory cortex respond best to low frequencies; neurons at the other respond best to high frequencies. There are multiple auditory areas (much like the multiple areas in the visual cortex), which can be distinguished anatomically and on the basis that they contain a complete "frequency map." The purpose of this frequency map (known as a tonotopic map) is unknown and is likely to reflect the fact that the sensory epithelium of the auditory system, the cochlea, is arranged according to sound frequency. The auditory cortex is involved in tasks such as identifying and segregating auditory "objects" and identifying the location of a sound in space. Human brain scans have indicated that a peripheral bit of this brain region is active when trying to identify pitch. Individual cells consistently get excited by sounds at specific frequencies, or multiples of that frequency. The primary auditory cortex is about the same as Brodmann areas 41 y 42. It lies in the posterior half of the superior temporal gyrus and also dives into the lateral sulcus as the transverse temporal gyri (also called Heschl's gyri).  Área de Brodmann 41 This area is also known as anterior transverse temporal area 41 (H). It is a subdivision of the cytoarchitecturally-defined temporal region of cerebral cortex, occupying the anterior transverse temporal gyrus (H) in the bank of the lateral sulcus on the dorsal surface of the temporal lobe. Área de Brodmann 41 is bounded medially by the parainsular area 52 (H) and laterally by the posterior transverse temporal area 42 (H) (Brodmann-1909).  Área de Brodmann 42 This area is also known as posterior transverse temporal area 42 (H). It is a subdivision of the cytoarchitecturally-defined temporal region of cerebral cortex, located in the bank of the lateral sulcus on the dorsal surface of the temporal lobe. Área de Brodmann 42 is bounded medially by the anterior transverse temporal area 41 (H) and laterally by the superior temporal area 22 (Brodmann-1909).  Relationship to auditory system Areas of localization on lateral surface of hemisphere. Motor area in red. Area of general sensations in blue. Auditory area in green. Visual area in yellow. The psychic portions are in lighter tints. The auditory cortex is the most highly organized processing unit of sound in the brain. This cortex area is the neural crux of hearing, y, in humans, language and music. The auditory cortex is divided into three separate parts, the primary, secondary and tertiary auditory cortex. These structures are formed concentrically around one another, with the primary AC in the middle and the tertiary AC on the outside. The primary auditory cortex is tonotopically organized, which means that certain cells in the auditory cortex are sensitive to specific frequencies. This is a fascinating function which has been preserved throughout most of the audition circuit. This area of the brain “is thought to identify the fundamental elements of music, such as pitch and loudness (17).” This makes sense as this is the area which receives direct input from the medial geniculate nucleus of the thalamus. The secondary auditory cortex has been indicated in the processing of “harmonic, melodic and rhythmic patterns (17).” The tertiary auditory cortex supposedly integrates everything into the overall experience of music (17).  An evoked response study of congenitally deaf kittens by Klinke et al. utilized field potentials to measure cortical plasticity in the auditory cortex. These kittens were stimulated and measured against a control or un-stimulated CDC and normal hearing cats. The field potentials measured for artificially stimulated CDC was eventually much stronger than that of a normal hearing cat(8). This is in concordance with Eckart Altenmuller’s study where it was observed that students who received musical instruction had greater cortical activation than those who did not (9).  The auditory cortex exhibits some strange behavior pertaining to the gamma wave frequency. When subjects are exposed to three or four cycles of a 40 hertz click, an abnormal spike appears in the EEG data, which is not present for other stimuli. The spike in neuronal activity correlating to this frequency is not restrained to the tonotopic organization of the auditory cortex. It has been theorized that this is a “resonant frequencyof certain areas of the brain, and appears to affect the visual cortex as well (10).  Gamma band activation (20 Para 40 Hz) has been shown to be present during the perception of sensory events and the process of recognition. Kneif et al, in their 2000 estudiar, presented subjects with eight musical notes to well known tunes, such as Yankee Doodle and Frere Jacques. Randomly, the sixth and seventh notes were omitted and an electroencephalogram, as well as a magnetoencephalogram were each employed to measure the neural results. Específicamente, the presence of gamma waves, induced by the auditory task at hand, were measured from the temples of the subjects. The OSP response, or omitted stimulus response, was located in a slightly different position; 7 mm more anterior, 13 mm more medial and 13 mm more superior in respect to the complete sets. The OSP recordings were also characteristically lower in gamma waves, as compared to the complete musical set. The evoked responses during the sixth and seventh omitted notes are assumed to be imagined, and were characteristically different, especially in the right hemisphere (12). The right auditory cortex has long been shown to be more sensitive to tonality, while the left auditory cortex has been shown to be more sensitive to minute sequential differences in sound specifically speech. Hallucinations have been shown to produce oscillations which are parallel (although not exactly the same as) the gamma frequency range. Sperling showed in his 2004 study that auditory hallucinations produce band wavelengths in the range of 12.5-30 Hz. The bands occurred in the left auditory cortex of a schizophrenic and were controlled against 13 controls (18) . This aligns with the studies of people remembering a song in their minds; they do not perceive any sound, but experience the melody, rhythm and overall experience of sound. When schizophrenics experience hallucinations, it is the primary auditory cortex which becomes active. This is characteristically different from remembering a sound stimulus, which only faintly activates the tertiary auditory cortex (16). By deduction, an artificial stimulation of the primary auditory cortex should elicit an incredibly real auditory hallucination. The termination of all audition and music into the tertiary auditory cortex creates a fascinating nexus of aural information. If this theory is true, it would be interesting to study a subject with a damaged, TAC or one with artificially suppressed function. This would be very difficult to do as the tertiary cortex is simply a ring around the secondary, which is a ring around the primary AC. Tone is perceived in more places than just the auditory cortex; one specifically fascinating area is the rostromedial prefrontal cortex (13). Janata et al, in their 2002 estudiar, used an fMRI machine to study the areas of the brain which were active during tonality processing. The result of which displayed several areas which are not normally considered to be part of the audition process. The rostromedial prefrontal cortex is a subsection of the medial prefrontal cortex, which projects to the amygdala, and is thought to aid in the inhibition of negative emotion (14). The medial prefrontal cortex is thought to be the core developmental difference between the impulsive teenager and the calm adult. The rostromedial prefrontal cortex is tonality sensitive, meaning it is activated by the tones and frequencies of resonant sounds and music. It could be hypothesized that this is the mechanism by which music ameliorates the soul (o, if one prefers, the limbic system).  Referencias 8) Klinke, Rainer. Recruitment of the auditory cortex in congenitally deaf cats by long-term cochlear electrostimulation. Science v. 285 no. 5434 (Septiembre 10 1999) p. 1729-33 9) Strickland. Music and the brain in childhood development. Childhood Education, Invierno 2001 v78 i2 p100(4) 10) Bertrand, et al. Object Representation and Gamma Oscillations. HTTP://biomag2000.hut.fi/papers_audition.html 11) HTTP://ist-socrates.berkeley.edu/~jmp/LO2/6.html 12) Knief, A et al. Oscillatory Gamma band and Slow brain Activity Evoked by Real and Imaginary Musical Stimuli. HTTP://biomag2000.hut.fi/papers_audition.html 13) Petr Janata et al. The Cortical Topography of Tonal Structures Underlying Western Music. Ciencia, Para. 298, Emitir 5601, 2167-2170 , 13 Diciembre 2002 14) Cassel, Topography of projections from the medial prefrontal cortex to the amygdala in the rat. Brain Res Bull. 1986 Sep;17(3):321-33 15) Cant NB, Parallel auditory pathways: projection patterns of the different neuronal populations in the dorsal and ventral cochlear nuclei. Brain Res Bull. 2003 Jun 15;60(5-6):457-74.  16) Abbott, Alison Music, maestro, please! Nature v. 416 no. 6876 (Marzo 7 2002) 17) Ferragamo MJ. Octopus cells of the mammalian ventral cochlear nucleus sense the rate of depolarization. J Neurofisiol. 2002 Mayo;87(5):2262-70 External links BrainInfo at the University of Washington ancil-77 - area 41 BrainInfo at the University of Washington ancil-78 - area 42 BrainMaps at UCDavis primary%20auditory%20cortex http://www.al-shia.com/html/eng/p.php?p=quran&url=Al-Quran See also Auditory system Brodmann area Noise health effects v·d·e Sensory system: Sistemas auditivo y vestibular (AT A15.3, Georgia 10.1029) Pabellón auricular externo (Hélice, antihélix, trago, antitrago, Incisión anterior de la oreja, Lóbulo de oreja) • Canal auditivo• Auricular muscles Eardrum (Forma, parte flácida) Oído medio Cavidad timpánica Pared laberíntica/medial: ventana ovalada· Round windowSecondary tympanic membraneProminence of facial canalPromontory of tympanic cavity Membranous wall/lateral Mastoid wall/posterior: Mastoid cellsAditus to mastoid antrumPyramidal eminence Carotid wall/anterior Tegmental wall/roof: Receso epitimpánico Pared/suelo yugular Osículos Martillo (cuello de martillo, Ligamento superior del martillo, Ligamento lateral del martillo, Ligamento anterior del martillo) · Incus (Ligamento superior del yunque, Ligamento posterior del yunque) · estribo (Ligamento anular del estribo) Muscles Stapedius· Tensor tympani Eustachian tube Bony part of pharyngotympanic tube· Cartilage of pharyngotympanic tube (gaitero toro) Oído interno/ (laberinto membranoso, laberinto óseo) Auditory system Cochlear labyrinth General cochlea Scala vestibuliHelicotremaScala tympaniModiolusCochlear cupula Perilymphatic space PerilymphCochlear aqueduct Cochlear duct / scala media Reissner's/vestibular membraneBasilar membrane EndolymphStria vascularisSpiral ligament Organ of Corti: StereociliaTectorial membraneSulcus spiralis (externo, interno) • Spiral limbus Vestibular system/ Vestibular labyrinth Static/translations/vestibule/endolymphatic duct: utrículo (Mancha) · Saccule (Mancha, saco endolinfatico) · Kinocilium· OtolithVestibular aqueductCanalis reuniens Kinetic/rotations: Canales semicirculares (Superior, Posterior, Horizontal) • Ampullary cupulaAmpullae (Crista ampullaris) {| clase="navbox plegable nowraplinks" estilo="margen:auto; " [. v]·[. d]·[. e] M: Oído anat (e/p)/física/desarrollo noco/cong, epón proc, droga (S2) |} telencéfalo (telencéfalo, corteza cerebral, hemisferios cerebrales) - editar surcos/fisuras primarias: longitudinal medial, lateral, central, parietoöccipital, calcarina, lóbulo frontal cingulado: giro precentral (corteza motora primaria, 4), surco precentral, giro frontal superior (6, 8), giro frontal medio (46), giro frontal inferior (área de Broca, 44-pars opercularis, 45-pars triangularis), corteza prefrontal (corteza orbitofrontal, 9, 10, 11, 12, 47) lóbulo parietal: surco postcentral, giro postcentral (1, 2, 3, 43), lóbulo parietal superior (5), lóbulo parietal inferior (39-giro angular, 40), precúneo (7), Lóbulo occipital del surco intraparietal: corteza visual primaria (17), cuneus, giro lingual, 18, 19 (18 y 19 abarcar todo el lóbulo) lóbulo temporal: giro temporal transversal (41-42-corteza auditiva primaria), giro temporal superior (38, 22-el área de Wernicke), giro temporal medio (21), giro temporal inferior (20), giro fusiforme (36, 37) lóbulo límbico/giro fornicado: corteza cingulada/giro cingulado, cingulado anterior (24, 32, 33), cingulado posterior (23, 31), istmo (26, 29, 30), giro parahipocampal (corteza piriforme, 25, 27, 35), corteza entorrinal (28, 34) corteza subcortical/insular: rhinencephalon, bulbo olfativo, cuerpo calloso, ventrículos laterales, septum pellucidum, epéndimo, cápsula interna, corona radiata, Formación del hipocampo de la cápsula externa.: giro dentado, hipocampo, subículo ganglios basales: Estriado (núcleo caudado, Putamen), núcleo lentiforme (Putamen, globus pallidus), claustrum, cápsula extrema, amígdala, núcleo accumbens Algunas categorizaciones son aproximaciones, and some Brodmann areas span gyri. 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